ISI PUBLICATIONS (Google Scholar metrics)


Liew T-S & Clements GR (2020). Whittenia, a new genus of land snails from Perak, Peninsular Malaysia (Gastropoda:Diplommatinidae). RAFFLES BULLETIN OF ZOOLOGY Supplement 35: 143-148. PDF

Aziz SA, Chua MAH & Clements GR (2020). Catch and release: Novel predation strategy by white-bellied sea-eagle (Haliaeetus leucogaster) on island flying fox (Pteropus hypomelanus) demonstrates flying fox swimming ability. JOURNAL OF BAT RESEARCH AND CONSERVATION12: 64-66. PDF


Macdonald DW, Bothwell HM, Kaszta Z, Ashl E, Bolongon G, Burnham D, Can OE, Campos‐Arceiz A, Channa P, Clements GR, Hearn AJ, Hedges L, Htun S, Kamler JF, Kawanishi K, Macdonald EA, Mohamad SW, Moore J, Naing H, Onuma M, Penjor U, Rasphone A, Rayan DM, Ross J, Singh P, Tan CKW, Wadey J, Yadav BP & Cushman SA (2019). Multi‐scale habitat modelling identifies spatial conservation priorities for mainland clouded leopards (Neofelis nebulosa). DIVERSITY AND DISTRIBUTIONS 25: 1639-1654. DOI

Pardo LE, Campbell MJ, Cove MV, Edwards W, Clements GR & Laurance WF (2019). Land management strategies can increase oil palm plantation use by some terrestrial mammals in Colombia. SCIENTIFIC REPORTS 9: 7812. DOI

Coleman J, Ascher JS, Bickford D, Buchori D, Cabanban A, Chisholm RA, Chong KY, Christie P, Clements GR, dela Cruz TEE, Dressler W, Edwards DP, Francis CM, Friess DA, Giam X, Gibson L, Huang D, Hughes AC, Jaafar Z, Koh LP, Kudavidanage EP, Lee B, Lee J, Lee TM, Leggett M, Leimona B, Linkie M, Luskin M, Lynam A, Meijaard E, Nijman V, Olsson A, Page S, Parolin P, Peh, KS-H, Posa MRC, Prescott GW, Rahman SA, Ramchunder SJ, Rao M, Reed J, Richards DR, Slade E, Steinmetz R, Tan PY, Todd PA, Vo ST, Webb EL, Ziegler AD & Carrasco LR (2019). Top 100 questions for biodiversity conservation in Southeast Asia. BIOLOGICAL CONSERVATION 234: 211-220. DOI – PDF


Clements GR, Aziz SA, Giam X, Bentrupperbaunmer J, Goosem M, Laurance S & Laurance WF (2018). Not everyone wants roads: assessing indigenous people’s support for roads in a globally important tiger conservation landscape. HUMAN ECOLOGY 46: 909-915. DOI – PDF

Liang SH, Yong DL, Abdul Kadir AH, Pazil AP, Rahmah I, Hazril AH, Ahmad Khusaini MKS, Trai LT & Clements GR (2018). Peninsular Malaysia’s forgotten pheasant: recent records and distribution of the Crested Argus Rheinardia ocellata. FORKTAIL 34: 48-51.

Wijedasa L, Sloan S, Page SE, Clements GR, Lupascu M & Evans TA (2018). Carbon emissions from Southeast Asian peatlands will increase despite REDD+ program. GLOBAL CHANGE BIOLOGY 24: 4598-4613. DOI – PDF

Pardo-Vargas LE, Campbell MJ, Edwards W, Clements GR & Laurance WF (2018). Terrestrial mammal responses to oil palm dominated landscapes in Colombia. PLOS ONE 13(5): e0197539. DOI – PDF

Ratnayeke S, van Manen FT, Clements GR, Mohd Kulaimi NA & Sharp SP (2018). Carnivore hotspots in Peninsular Malaysia and their landscape attributes. PLOS ONE 13(4): e0194217. DOI – PDF


Alamgir M, Campbell MJ, Sloan S, Goosem M, Clements GR, Mahmoud IM & Laurance WF (2017). Economic, socio-political and environmental risks of road development in the tropics. CURRENT BIOLOGY 27: R1130-R1140. DOI – PDF

Aziz SA, Clements GR, McConkey KR, Sritongchuay T, Pathil S, Abu Yazid MNH, Campos-Arceiz A, Forget P-M & Bumrungsri S (2017). Pollination by the locally endangered island flying fox (Pteropus hypomelanus) enhances fruit production of the economically important durian (Durio zibethinus). ECOLOGY AND EVOLUTION 7: 8670-8684 (Cover article). DOI – PDF

Scotson L, Johnston LR, Iannarillil F, Wearn OR, Mohd-Azlan J, Wong WM, Gray TNE, Dinata Y, Suzuki A, Willard CE, Frechette J, Loken B, Steinmetz R, Moßbrucker, AM, Clements GR & Fieberg J (2017). Best practices and software for the management and sharing of camera trap data for small and large scales studies. REMOTE SENSING IN ECOLOGY AND EVOLUTION 3: 158-172. DOI – PDF

Foon JK, Clements GR, Liew T-S (2017). Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZOOKEYS 682: 1-94. DOI – PDF

Aziz SA, Clements GR, Giam X, Forget P-M & Campos-Arceiz A (2017). Coexistence and conflict between the island flying fox (Pteropus hypomelanus) and humans on Tioman island, Peninsular Malaysia. HUMAN ECOLOGY 45: 377-389. PDFERRATUM

Aziz SA, Clements GR, Peng LY, Campos-Arceiz A, McConkey KR, Forget P-M & Gan HM (2017). Elucidating the diet and foraging ecology of the island flying fox (Pteropus hypomelanus) in Peninsular Malaysia through Illumina Next Generation Sequencing. PEERJ 5: e3176. PDF

Tan CKW, Rocha DG, Clements GR, Brenes-Mora E, Hedges L, Kawanishi K, Mohamad SW, Rayan DM, Bolongon G, Moore J, Wadey J, Campos-Arceiz, A & Macdonald DW (2017). Habitat use and predicted range of the mainland clouded leopard Neofelis nebulosa in Peninsular Malaysia. BIOLOGICAL CONSERVATION 206: 65-74.  DOIPDF

Fang F, Nguyen TH, Pickles R, Lam WY, Clements GR, An B, Singh A, Schwedock BC, Tambe M & Lemieux A (2017). PAWS-A Deployed Game-Theoretic Application to Combat Poaching. AI MAGAZINE 38: 23.  DOIPDF

Wijedasa LS, Jauhiainen J, Könönen M, Lampela M, Vasander H, LeBlanc M-C, Evers S, Smith TEL, Yule CM, Varkkey H, Lupascu M, Parish F, Singleton I, Clements GR, Aziz SA, Harrison ME, Cheyne S, Anshari GZ, Meijaard E, Goldstein JE, Waldron S, Hergoualc’h K, Dommain R, Frolking S, Evans CD, Posa MRC, Glaser PH, Suryadiputra N, Lubis R, Santika T, Padfield R, Kurnianto S, Hadisiswoyo P, Lim TM, Page SE, Gauci V, van der Meer PJ, Buckland H, Garnier F, Samuel MK, LiM LZ, Choo K, O’Reilly P, Warren M, Suksuwan S, Sumarga R, Jain A, Laurance WF, Couwenberg J, Joosten H, Vernimmen R, Hooijer A, Malins C, Cochrane MA, Perumal B, Siegert F, Peh K S-H, Comeau L-P, Verchot L, Harvey CF, Cobb A, Jaafar1 Z, Wösten H, Manuri S, Müller M, Giesen W, Phelps J, Yong DL, Silvius M, Wedeux BMM, Hoyt A, Osaki M, Takashi H, Takahashi H, Kohyama TS, Haraguchi A, Nugroho NP, Coomes DA, Quoi LP, Dohong A, Gunawan H, Gaveau DLA, Langner A, Lim FKS, Edwards DP, Giam X, van der Werf G, Carmenta R, Verwer CC, Gibson L, Grandois L, Graham LLB, Regalino J, Wich SA, Rieley J, Kettridge N, Brown C, Pirard R, Moore S, Capilla BR, Ballhorn U, Ho HC, Hoscilo A, Lohberger S, Evans TA, Yulianti N, Blackham G, Onrizal, Husson S, Murdiyarso D, Pangala S, Cole LES, Tacconi L, Segah H, Tonoto P, Lee JSH, Schmilewski G, Wulffraat S, Putra EI, Cattau ME, Clymo RS, Morrison R, Mujahid A, Miettinen J, Liew SC, Valpola S, Wilson D, D’Arcy L, Gerding M, Sundari S, Thornton SA, Kalisz B, Chapman SJ, Su ASM, Basuki I, Itoh M, Traeholt C, Sloan S, Sayok AK & Andersen R (2017). Denial of long-term issues with agriculture on tropical peatlands will have devastating consequences. GLOBAL CHANGE BIOLOGY 23: 977-982. DOI – PDF


Yaap B, Magrach A, Clements GR, McClure CJW, Paoli GD & Laurance WF. Large mammal use of linear remnant forests in an industrial pulpwood plantation in Sumatra, Indonesia (2016). TROPICAL CONSERVATION SCIENCE 9:1940082916683523. PDF

Nagulendran K, Padfield R, Aziz SA, Amir AA, Rahman ARA, Latiff MA, Zafir Ahmad AW, Aida GQ, Tan A, Arifah S, Awang N, Azhar N, Balu P, Gan PC, Ning H, Sharma DK, Lavanya RI, Lim T, Mahendra S, Rayan DM, McGowan S, Paxton M, Mohamed Z, Imam HRM, Abdullah MT, Ibrahim NAN, Puan CL, Rahimatsah A, Clements GR, Mohamed ISM, Saw LG, Shashi K, Sivananthan E, Surin S, Vanitha P, Wadey J, Hasmah WMW, Wong EP, Wong PM, Yeap CA & Campos-Arceiz A. (2016). A multi-stakeholder strategy to identify conservation priorities in Peninsular Malaysia. COGENT ENVIRONMENTAL SCIENCE 2: 1254078. PDF 

Lee P-S, Gan HM, Clements GR and Wilson J-J (2016). Field calibration of blowfly-derived DNA against traditional methods for assessing mammal diversity in tropical forests. GENOME 59: 1008-2022. DOI – PDF 

Brodie JF, Paxton M, Nagulendran K, Balamurugan G, Clements GR, Reynolds G, Jain A and Hon J (2016). Connecting science, policy, and implementation for landscape-scale habitat connectivity. CONSERVATION BIOLOGY 30: 950-961. DOIPDF

Rostro-García S, Kamler J, Ash E, Clements GR, Gibson L, Lynam AJ, McEwing R, Naing H and Paglia S (2016). Endangered leopards: range collapse of the Indochinese leopard (Panthera pardus delacouri) in Southeast Asia. BIOLOGICAL CONSERVATION 201: 293-300. DOIPDF 

Liew T-S, Price L, Clements GR (2016). Using Google Earth to improve management of threatened limestone karst ecosystems in Peninsular Malaysia. TROPICAL CONSERVATION SCIENCE 9: 903-920. PDF 


Hedges L, Morrant DS, Campos-Arceiz A and Clements GR (2015). Feasibility of using scent-baited hair traps to monitor carnivore populations in Peninsular Malaysia. TROPICAL CONSERVATION SCIENCE 8: 975-982. PDF

Wijedasa L, Posa MRC and Clements GR (2015). Peat fires: consumers to help beat them out. NATURE 527:305. DOIPDF

Vargas LEP, Laurance WF, Clements GR and Edwards W (2015). The impacts of oil palm agriculture on Colombia’s biodiversity: what we know and still need to know. TROPICAL CONSERVATION SCIENCE 8: 828-845. PDF

Hedges L, Lam WY, Campos-Arceiz A, Mark Rayan D, Laurance WF, Latham CJ, Saaban S and Clements GR (2015), Melanistic leopards reveal their spots: infrared cameras provide a population-density estimate of leopards in Malaysia. JOURNAL OF WILDLIFE MANAGEMENT 79: 846-853. DOIPDF


Clements GR, Lynam, AJ, Gaveau D, Yap WL, Lhota S, Goosem M, Laurance S & Laurance WF. (2014) Where and how are roads endangering mammals in Southeast Asia’s forests? PLOS ONE 9: e115376. DOIPDF

Laurance WF, Clements GR, Sloan SP, O’Connell CO, Mueller NP, Goosem M, Venter O, Edwards DP, Phelan B, Balmford A, van Der Ree R and Arrea IB (2014). A global strategy for road building. NATURE 513: 229-232. DOIPDF

Jain A, Kwek YC, Chua MAH and Clements GR (2014). Moving away from paper corridors in Southeast Asia. CONSERVATION BIOLOGY 28: 889-891. DOIPDF


Kawanishi K, Clements GR, Gumal M, Goldthrope G, Yasak MN and Sharma DSK (2013). Using BAD for good: how best available data facilitated a precautionary policy change to better protect tiger prey in Peninsular Malaysia. ORYX 47: 420-426. DOIPDF

Linkie M, Guillera-Arroita G, Smith J, Ario A, Bertagnolio G, Cheong F, Clements GR, Dinata Y, Duangchantrasiri S, Fredriksson G, Gumal MT, Horng LK, Kawanishi K, Khakim FR, Kiswayadi D, Lubis AH, Lynam AJ, Maryati M, Maung M, Ngoprasert D, Novarino W, O’Brien TGO, Parakkasi K, Peters H, Priatna D, Rayan DM, Seuaturien N, Shwe NM, Steinmetz R, Sugesti AM, Sunarto S, Sunquist ME, Umponjan M, Wibisono HT, Wong CCT and Zulfahmi (2013). Conserving cryptic mammals on camera: assessing the utility of range wide camera trap data for conserving the endangered Asian tapir. BIOLOGICAL CONSERVATION 162: 107-115. DOIPDF

Aziz SA, Clements GR, Rayan DM and Sankar P (2013). Why conservationists should be concerned about natural resource legislation affecting indigenous peoples’ rights: Lessons from Peninsular Malaysia. BIODIVERSITY AND CONSERVATION 22: 639-656. DOIPDF

Marzuki ME and Clements GR (2013). A new species and record of cyclophorid snail (Mollusca: Prosobranchia) from Terengganu, Peninsular Malaysia. RAFFLES BULLETIN OF ZOOLOGY 61: 21-24. PDF

Lim T-LN, Giam X, Byrnes, G and Clements GR (2013). Occurrence of the Sunda colugo (Galeopterus variegatus) in the tropical forests of Singapore: A Bayesian approach. MAMMALIAN BIOLOGY 78: 63-67. DOI – PDF


Clements GR, Rayan DM, Aziz SA, Kawanishi K, Traeholt C, Magintan D, Yazi MFA, and Tingley R (2012). Predicting the distribution of the Asian tapir (Tapirus indicus) in Peninsular Malaysia using maximum entropy modelling. INTEGRATIVE ZOOLOGY 7: 402-409. DOI – PDF

Rayan DM, Shariff M, Doward L, Aziz S, Clements GR, Wong C, Traeholt C and Magintan D (2012). Estimating the population density of the Asian tapir Tapirus indicus from a selectively logged forest in Peninsular Malaysia. INTEGRATIVE ZOOLOGY 7: 373-380.  DOI – PDF

Wijedasa LS, Sloan S, Michelakis DG and Clements GR (2012). Overcoming limitations with Landsat imagery for mapping of peat swamp forests in Sundaland. REMOTE SENSING 4: 2595-2618. DOI – PDF 

Clements GR and Tan SK (2012). A new genus and species of camaenid snail (Mollusca: Pulmonata) from Terengganu, Peninsular Malaysia. RAFFLES BULLETIN OF ZOOLOGY Supplement no. 25: 259-263. PDF


Giam X, Clements GR, Aziz SA, Chong KY and Miettinen J (2011). Rethinking the ‘back to wilderness’ concept for Sundaland’s forests. BIOLOGICAL CONSERVATION 144: 3149-3152. DOI – PDF

Bradshaw CJA,Clements GR, Brook BW and Laurance WF (2011). Better SAFE than sorry. FRONTIERS IN ECOLOGY AND THE ENVIRONMENT 9: 487-488. DOI – PDF

Ahmad Zafir AW, Payne J, Azlan M, Lau CF, Sharma DSK, Alfred R, Williams AC, Nathan S, Ramono WS and Clements GR (2011). Now or never: what will it take to save the Sumatran rhinoceros (Dicerorhinus sumatrensis) from extinction? ORYX 45: 225-233. DOI – PDF

Clements GR, Bradshaw CJA, Brook BW and Laurance WF (2011). The SAFE index: using a threshold population target to measure relative species threat. FRONTIERS IN ECOLOGY AND THE ENVIRONMENT 9: 521-525. DOI – PDF

Laurance WF, Kakul T, Keenan RJ, Sayer J, Clements GR, Villegas F and Sodhi NS (2011). Predatory corporations, failing governance and the fate of forests in Papua New Guniea. CONSERVATION LETTERS 4: 95-100. DOI – PDF

Tan SK, and Clements GR (2011). Limicolaria flammea: another potentially invasive African land snail in tropical Asia. TROPICAL CONSERVATION SCIENCE 4: 97-102. PDF

Peh K S-H, Soh MK, Sodhi NS, Laurance WF, Ong DJ and Clements GR (2011). Up in the clouds: is sustainable use of tropical montane cloud forests possible in Malaysia? BIOSCIENCE 61: 27-38. DOI – PDF


Clements GR, Sayer J, Boedhihartono AK, Venter O, Lovejoy T, Koh LP and Laurance WF (2010). Cautious optimism over Norway-Indonesia REDD pact. CONSERVATION BIOLOGY 24: 1437-1438. DOI – PDF

Clements GR, Giam X and Laurance WF (2010). Will India’s mega-census leave a mega-footprint? CURRENT SCIENCE 99: 556. PDF

Aziz SA, Laurance W and Clements R (2010). Forests reserved for rubber? FRONTIERS IN ECOLOGY AND THE ENVIRONMENT 8: 178. DOI PDF

Sodhi NS, Koh LP, Clements R, Wanger TC, Hill JK, Clough Y, Tscharntke T, Posa MRC and Lee TM (2010). Conserving Southeast Asian forest biodiversity in human-modified landscapes. BIOLOGICAL CONSERVATION 143: 2375-2384. DOI PDF

Clements R, Rayan DM, Ahmad Zafir AW, Venkataraman A, Alfred R, Payne J, Ambu L and Sharma DSK (2010). Trio under threat: can we secure the future of rhinos, elephants and tigers in Malaysia? BIODIVERSITY AND CONSERVATION 19: 1115-1136. DOI – PDF


Clements R, Foo R, Othman S, Rahman U, Syed Mustafa SR and Zulkifli R (2009). Islam, turtle conservation and coastal communities. CONSERVATION BIOLOGY 23: 516-517. DOI – PDF


Clements R, Lu XX, Ambu S, Schilthuizen M and Bradshaw C (2008). Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts. BIOLOGICAL CONSERVATION 141: 2751-2764. DOI – PDF

Tan SK and Clements R (2008). Taxonomy and distribution of the Neritidae (Mollusca: Gastropoda) in Singapore. ZOOLOGICAL STUDIES 47: 481-494. PDF

Clements R, Liew T-S, Vermeulen JJ and Schilthuizen M (2008). Further twists in gastropod shell evolution. BIOLOGY LETTERS 4: 179-182. DOI PDF


Clements R and Posa MRC (2007). Conservationists could slip up in oil-palm enterprise. NATURE 449: 443. DOI PDF

Clements R, Bickford D and Lohman D (2007). Can YouTube save the planet? THE SCIENTIST 21: 26. PDF


Clements R, Sodhi NS, Ng PKL and Schilthuizen M (2006). Limestone karsts of Southeast Asia: imperiled arks of biodiversity. BIOSCIENCE 56: 733-742 (Cover article). DOI PDF

Clements R, Koh LP, Lee TM, Meier R and Li D (2006). Importance of reservoirs for the conservation of freshwater molluscs in a tropical urban landscape. BIOLOGICAL CONSERVATION 128: 136-146. DOI – PDF

Clements R (2006). Oophana tiomanensis, a new streptaxid (Gastropoda: Pulmonata: Streptaxidae) from Pulau Tioman, Peninsular Malaysia. RAFFLES BULLETIN OF ZOOLOGY 54: 125-127. PDF


Clements R and Li D (2005). Regulation and non-toxicity of the spit from the pale spitting spider Scytodes pallida (Araneae: Scytotidae). ETHOLOGY 111: 311-321. DOI PDF


Tan SK, Chan S-Y and Clements GR. 2012. A Guide to Snails and other Non-marine Molluscs of Singapore. Science Centre, Singapore. 176pp. ISBN: 978-981-08-9549-5. Book Cover


de Gouvenain, R and Clements GR (2014). Ecological and Evolutionary Processes. In: Encyclopedia of Natural Resources. Yeqiao YQ (editor). pp. 138-142. Taylor & Francis Group. DOI

Koh LP, Kettle CJ, Sheil D, Lee TM, Giam X, Gibson L and Clements GR. Biodiversity state and trends in Southeast Asia (2013). In: Encyclopedia of Biodiversity, 2nd edition. Levin S (editor). pp 509-527. Elsevier Inc. ISBN: 978-0-12-384720-1. DOI PDF


Fang F, Nguyen TH, Pickles R, Lam WY, Clements GR, An A, Singh A, Tambe M & Lemieux A (2016). Deploying PAWS: Field Optimization of the Protection Assistant for Wildlife Security. In  PROCEEDINGS OF THE INNOVATIVE APPLICATIONS OF ARTIFICIAL INTELLIGENCE (Winner of Deployed Application Award). PDF

Foon JK, Tan SK & Clements GR (2015). A new species of Kenyirus (Pulmonata, Camaenidae) from Peninsular Malaysia. JOURNAL OF TROPICAL BIOLOGY AND CONSERVATION 12: 25-34. PDF

Rayan DM & Clements GR (2015). What we know and don’t know about the Malayan Tiger. MYCAT TRACKS 5: 18-19. PDF

Lam WY, Hedges L & Clements GR (2014). First record of a clouded leopard predating on a binturong. CAT NEWS 60: 16. PDF

Wijedasa L, Tan SK, Clements GR & Sng D (2014). Three snails in a slugfest. NATURE WATCH 22: 2-5.

Hedges L, Clements GR, Aziz SA, Yap W, Laurance S, Goosem M & Laurance WF (2013). The status of small mammalian carnivores in a threatened wildlife corridor in Peninsular Malaysia. SMALL CARNIVORE CONSERVATION 49: 9-14. PDF

Wijedasa L, Aziz SA, Campos-Arceiz A and Clements GR (2013). Does social media “like” conservation? THE SCIENTIST PDF

Clements GR and Tan SK (2012). Kenyirus sodhii: a tribute to a special place and conservation scientist. MALAYSIAN NATURALIST 66: 16-17. PDF

Clements GR, Yap W and Henry P (2012). Towards safer passages: the Kenyir wildlife corridor project. MALAYSIAN NATURALIST 65: 56-59. PDF

Laurance WF and Clements R (2010). How do roads affect imperilled mammals and indigenous people in Malaysia? SOLITAIRE 21: 10-11. PDF

Liew T-S, Clements R and Schilthuizen M (2008). Sampling micromolluscs in tropical forests: one size does not fit all. ZOOSYMPOSIA 1: 271-280. PDF

Vermeulen JJ and Clements R (2008). Another twist in the tale: a new species of Opisthostoma (Gastropoda, Diplommatinidae) from Peninsular Malaysia. BASTERIA 72: 263-268. PDF

Schilthuizen M and Clements R (2008). Tracking land snail extinctions from space. TENTACLE 16: 8-9 (IUCN/SSC Mollusc Specialist Group Newsletter). PDF



Clements R, Gumal M, Krishnasamy K, Rayan M, Shepherd C & Lynam A. 2015. In: Kawanishi K 2015. Panthera tigris jacksoni. IUCN Red List of Threatened Species. Version 2015-4. 

Whitten T, Clements R & Price L. 2013. Liphistius kanthan. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1. 

Clements R 2009. Opisthostoma michaelis. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1.

Clements R 2009. Opisthostoma retrovertens. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1.

Clements R 2009. Opisthostoma sciaphilum. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1.

Clements R 2009. Opisthostoma senex. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1.

Clements R 2009. Opisthostoma thersites. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1.

Clements R 2009. Opisthostoma trapezium. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.1.


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